簡易檢索 / 詳目顯示

研究生: 陳順其
論文名稱: 墾丁國家公園臺灣梅花鹿活動模式及其對棲地之利用
指導教授: 王穎
學位類別: 博士
Doctor
系所名稱: 生命科學系
Department of Life Science
論文出版年: 2002
畢業學年度: 90
語文別: 中文
論文頁數: 130
中文關鍵詞: 台灣梅花鹿行為活動模式活動範圍食性磨痕啃剝樹皮
論文種類: 學術論文
相關次數: 點閱:313下載:9
分享至:
查詢本校圖書館目錄 查詢臺灣博碩士論文知識加值系統 勘誤回報
  • 為了解臺灣梅花鹿(Cervus nippon taiouanus)野放後之行為及棲地利用,自1994年7月至2000年8月在墾丁國家公園進行調查。個體活動以日間較高。其中位移以乾季較高,休息以溼季較高。雌性個體之位移較雄性高,反芻則反之。由無線電追蹤野放個體之活動量,顯示其平均日活動為57.2%,每天有4至6次高峰,雌雄相差不大。其活動受天候及人之影響,雨天較晴天低(45.7±23.2與62.9±24.7%);陰天有落山風時亦較晴天低(52.3±24.7%與74.7±18.1%)。非假日遊客較少時較假日高(57.1% vs. 43.6%)。就野放個體活動範圍而言,所有鹿隻之活動範圍(95%MCP)為755.2公頃。平均日活動範圍為12.26±8.59公頃,乾季之季平均活動範圍為101.3公頃,濕季為51.7公頃,年活動範圍平均為159.8公頃。就食性而言,共記錄51科157種鹿食植物,其中以喬木與草本居多。其中有8種喜食及86種進食種類。在224種被鹿、水牛及山羊取食的植物中,以羊的食性較廣(204種),鹿次之(157種),牛較少(68種),其中三者共食者有52種,重疊性為36.4%。鹿與羊相似度為76.3%,鹿與牛為49.3%,牛與羊為46.7%,顯示鹿與牛或羊間之食物重疊性較牛羊間為高。鹿對當地植被影響之結果顯示,樣區內、外植被無論有無鹿隻覓食均朝正向演替。另由一區調查結果顯示,底層植物之減少與鹿食喜好程度有關。就磨樹行為而言,鹿對部分樹種有所偏好(X2,p<0.005)。計有27科55種430棵植物被磨,其中有重複被磨,重複性達63.6%。被磨致死的樹種有14種30棵,佔穿越線所有棵數之0.2%。磨痕多寡與該區食源等級高低有關(ANOVA, p<0.05)。磨痕出現的時間與鹿隻之繁殖季節有關。就啃食樹皮行為而言,啃食樹皮僅在復育第四區發現,計有10科15種共1010棵被啃,顯示鹿對被啃樹種有選擇性。啃痕各月均有出現,數量以乾季末期及濕季初期較高。被啃致死的樹種共有9種427棵,死亡率為42.3%。其有啃痕之出現應與本區鹿群密度(每公頃高於2.1頭)較其他地區(每公頃低於1.8頭)高及棲地品質差有關。 From July 1994 to August 2000 field work had been conducted to study the behavior and habitat use of the Formosan sika deer (Cervus nippon taiouanus) in Kenting National Park. The result showed that deer had higher daytime activity than that of nighttime. Movement was higher in dry season; while rest was higher in the wet season. Female had higher activity in movement; and male had higher activity in rumination. The average daily activity was 57.2% among radio-tracked deer with 4 to 6 active peaks each day. There was no difference between sexes in activity. Weather and tourist also affected deer activity. Deer had lower activity in cloudy days with strong wind (52.3%±24.7%) than those in sunny days (74.7%±18.1%). Similar trend was found between rainy days (45.7%±23.2%) and sunny days (62.9%±24.7%). Deer had higher activity in weekdays than those in weekends and other holidays (57.1% vs. 43.6%). The average annual home range was 159.8 ha and daily range was 12.24±8.59 ha. The average dry season range was 101.3ha, and wet season range was 51.7ha. In terms of food habits, 157 species mainly woody and herbaceous plants from 51 families were recorded. Among those species, 8 were preferred and 86 were commonly used by the deer. Two hundred twenty four species were recorded to be used by 3 herbivores in the study area. Among those species, 204 were used by goat, 157 by deer, 68 by water buffalo, and 52 by all 3 herbivores. The similarities between any 2 different herbivores and among 3 herbivores were 76.3% between deer and goat, 49.3% between deer and buffalo, 46.7% between goat and buffalo, and 36.4% among all 3 species. Deer had a higher food overlap with the other 2 species. The effect of deer activity on plant succession was insignificant from the observation on the plant composition in the sample plots. However, the decrease of undercover in the forest floor was related to the food habit of the deer. Four hundred thirty plants from 27 families and 55 species were rubbed by the deer. Among those plants 63.6% had repeated damages and some species was preferred (X2 test, p< 0.005). Thirty plants from 14 species had fatal damage which composed of 0.2% of all trees surveyed in transect lines. There was significant difference between number of rubbed tree and food quantity in the sampling plots (ANOVA, p< 0.05). The occurrence of tree rubbing was coincided with rutting season. The bark stripping was found only in Sec. 4 in the enclosed area. Those 1010 plants from 10 families and 15 species were bark stripped in comparison with over 100 untouched species indicated that deer selected certain species to rub. The activity was recorded every month with the peak activity in the late dry and early wet season. Four hundred and twenty seven plants from 9 species had fatal injury. The mortality of stripped trees was 42.3%. The occurrence of bark stripping in Sec. 4 was probably due to high deer density (> 2.1 deer/ha) of the area than those of other areas (< 1.8 deer/ha) and poor habitat quality.

    From July 1994 to August 2000 field work had been conducted to study the
    behavior and habitat use of the Formosan sika deer (Cervus nippon taiouanus)
    in Kenting National Park. The result showed that deer had higher daytime
    activity than that of nighttime. Movement was higher in dry season; while
    rest was higher in the wet season. Female had higher activity in movement;
    and male had higher activity in rumination. The average daily activity was 57.
    2% among radio-tracked deer with 4 to 6 active peaks each day. There was no
    difference between sexes in activity.  Weather and tourist also affected
    deer activity. Deer had lower activity in cloudy days with strong wind (52.3%
    ±24.7%) than those in sunny days (74.7%±18.1%). Similar trend was found
    between rainy days (45.7%±23.2%) and sunny days (62.9%±24.7%). Deer had
    higher activity in weekdays than those in weekends and other holidays (57.1%
    vs. 43.6%). The average annual home range was 159.8 ha and daily range was 12.
    24±8.59 ha. The average dry season range was 101.3ha, and wet season range
    was 51.7ha. In terms of food habits, 157 species mainly woody and herbaceous
    plants from 51 families were recorded. Among those species, 8 were preferred
    and 86 were commonly used by the deer. Two hundred twenty four species were
    recorded to be used by 3 herbivores in the study area. Among those species,
    204 were used by goat, 157 by deer, 68 by water buffalo, and 52 by all 3
    herbivores. The similarities between any 2 different herbivores and among 3
    herbivores were 76.3% between deer and goat, 49.3% between deer and buffalo,
    46.7% between goat and buffalo, and 36.4% among all 3 species. Deer had a
    higher food overlap with the other 2 species. The effect of deer activity on
    plant succession was insignificant from the observation on the plant
    composition in the sample plots. However, the decrease of undercover in the
    forest floor was related to the food habit of the deer. Four hundred thirty
    plants from 27 families and 55 species were rubbed by the deer. Among those
    plants 63.6% had repeated damages and some species was preferred (X2 test, p<
    0.005). Thirty plants from 14 species had fatal damage which composed of 0.2%
    of all trees surveyed in transect lines. There was significant difference
    between number of rubbed tree and food quantity in the sampling plots (ANOVA,
    p< 0.05). The occurrence of tree rubbing was coincided with rutting season. 
    The bark stripping was found only in Sec. 4 in the enclosed area. Those 1010
    plants from 10 families and 15 species were bark stripped in comparison with
    over 100 untouched species indicated that deer selected certain species to
    rub. The activity was recorded every month with the peak activity in the late
    dry and early wet season. Four hundred and twenty seven plants from 9 species
    had fatal injury. The mortality of stripped trees was 42.3%. The occurrence
    of bark stripping in Sec. 4 was probably due to high deer density (> 2.1 deer/
    ha) of the area than those of other areas (< 1.8 deer/ha) and poor habitat

    中文摘要 ⅰ 英文摘要 ⅱ 壹、緒言 1 貳、研究地區 9 參、研究方法 10 一、個體研究行為 10 二、活動模式調查 10 三、活動範圍及空間分佈 12 四、棲地利用 14 五、台灣梅花鹿、水牛及山羊食性之調查 15 六、臺灣梅花鹿對當地植物之影響 16 肆、結果 19 一、野化個體行為 19 二、活動模式 20 三、活動範圍及空間分佈 22 四、棲地利用 24 五、臺灣梅花鹿、水牛及山羊之食性 26 六、臺灣梅花鹿對當地植被的影響 31 伍、討論 39 一、野化臺灣梅花鹿個體行為的探討 39 二、臺灣梅花鹿活動模式的探討 40 三、臺灣梅花鹿活動範圍及空間分布的探討 41 四、棲地利用 44 五、鹿與共域草食獸牛羊之食性 46 六、磨樹及啃剝樹皮對當地植物可能造成之影響 53 七、臺灣梅花鹿對植被環境之影響 59 陸、總結 62 參考文獻 64 表目次 表1. 全日蒐集之無線電訊號以每半小時不同取樣長度估算活動比例(%)之分佈    ------------------------------------------------------------------------- ---------------77 表2. 無線電定位鹿隻活動範圍中不同定位點數估算活動面積(公頃)之分佈    ------------------------------------------------------------------------- ---------------77 表3. 野化台灣梅花鹿個體之行為比例(%)    ------------------------------------------------------------------------- ---------------77 表4. 野放臺灣梅花鹿日活動模式中活動所佔比率(%)    ------------------------------------------------------------------------- ---------------78 表5. 野放臺灣梅花鹿日活動模式之活動高峰數    ------------------------------------------------------------------------- ---------------78 表6. 野放臺灣梅花鹿雌性個體之活動範圍(公頃)    ------------------------------------------------------------------------- ---------------79 表7. 野放臺灣梅花鹿雄性個體之活動範圍(公頃)    ------------------------------------------------------------------------- ---------------80 表8. 台灣梅花鹿休息場所及排遺在各植被環境出現之比例(%)    ------------------------------------------------------------------------- ---------------81 表9 . 台灣梅花鹿食用植物屬性之分佈    ------------------------------------------------------------------------- ---------------81 表10 . 台灣梅花鹿食用植物調查方法比較    ------------------------------------------------------------------------- ---------------81 表11. 復育4區穿越線上台灣梅花鹿食痕之記錄植種    ------------------------------------------------------------------------- ---------------82 表12. 植生樣圍籬內外植種及數量之分佈    ------------------------------------------------------------------------- ---------------83 表13. 復育1區調查樣區中之優勢植物及其所佔總數之比例    ------------------------------------------------------------------------- ---------------84 表14. 復育1區穿越線上植種被梅花鹿磨之概況    ------------------------------------------------------------------------- ---------------85 表15. 台灣梅花鹿啃剝樹皮棵數及所佔比率    ------------------------------------------------------------------------- ---------------86 表16. 復育1區穿越線上植株大小數量、被鹿磨數量及所佔比例(%)    ------------------------------------------------------------------------- ---------------86 表17. 台灣梅花鹿啃剝樹皮之啃痕特性    ------------------------------------------------------------------------- ---------------87 表18. 復育1區穿越線上被鹿磨主要樹種數量分布及受害概況    ------------------------------------------------------------------------- ---------------87 表19. 復育1區樹木被台灣梅花鹿磨之各類植型之比例    ------------------------------------------------------------------------- ---------------88 表20. 復育1區被台灣梅花鹿磨之植株在不同地形之分布    ------------------------------------------------------------------------- ---------------88 表21. 復育1區樹木被台灣梅花鹿磨之磨痕所在處之地形概況及磨痕方向    ------------------------------------------------------------------------- ---------------89 表22. 社頂地區穿越線上地表狀況與台灣梅花鹿磨痕及排遺量之分佈    ------------------------------------------------------------------------- ---------------89 圖目次 圖1. 墾丁臺灣梅花鹿研究地區之相對位置與樣線及無線電收訊點之分佈------------90 圖2. 恆春地區月平均溫度(℃)及降水量(mm) --------------------------------------- ---------91 圖3. 無線電訊號次數與梅花鹿活動比例之關係-------------------------------------- -------91 圖4 . S6b型無線電訊號次數與活動比例之關係-------------------------------------- --------92 圖5 . S9型無線電訊號次數與活動比例之關係--------------------------------------- ---------92 圖6. 梅花鹿每一回合反芻時間的分佈---------------------------------------------- ------------93 圖7. 梅花鹿個體主要行為發生後最常接連發生的行為比例(%)------------------------- -93 圖8 復育區野化台灣梅花鹿覓食和位移的關係--------------------------------------- -------94 圖9 復育區野化台灣梅花鹿日活動模式--------------------------------------------- ----------94 圖10. 野化臺灣梅花鹿靜立與靜臥行為之日變化------------------------------------- -------95 圖11. 野化臺灣梅花鹿反芻與攝食行為之日變化------------------------------------- -------95 圖12. 野化臺灣梅花鹿位移與修飾行為之日變化------------------------------------- -------96 圖13. 野化臺灣梅花鹿雌雄個體活動模式之日變化----------------------------------- ------96 圖14. 野放臺灣梅花鹿雌雄個體乾濕季之日活動範圍--------------------------------- ----97 圖15. 野放臺灣梅花鹿雌雄個體乾溼季之季活動範圍--------------------------------- ----97 圖16. 野放臺灣梅花鹿無線電定位之歷年分佈--------------------------------------- --------98 圖17. 野放臺灣梅花鹿活動範圍內草原及非草原之分佈------------------------------- ----99 圖18. 臺灣梅花鹿餵食試驗中對不同型態植種之喜好程度----------------------------- --100 圖19. 臺灣梅花鹿餵食植種在不同區域出現之頻度----------------------------------- -----100 圖20. 臺灣梅花鹿與共域牛羊食性之比較------------------------------------------- ---------101 圖21. 共域草食獸鹿牛羊取食各型態植種比例之分布--------------------------------- ----101 圖22. 社頂地區樹木被台灣梅花鹿磨之植株1公尺及磨痕處之樹圍數量的分佈----102 圖23. 社頂地區樹木被台灣梅花鹿磨之磨痕在不同樣區出現之月變化----------------102 圖24. 社頂地區臺灣梅花鹿啃剝樹皮棵數之月變化----------------------------------- -----103 圖25. 復育區外台灣梅花鹿磨痕在不同食源等級植被樣站之分布---------------------- 103 附錄目次 附錄1. 研究期間及調查人力之分布------------------------------------------------ ---104 附錄2. 社頂地區野化個體各時刻之活動比例(%)----------------------------------105 附錄3. 社頂地區無線電訊號測得野放個體各時刻之活動比例(%)-------------106 附錄4. 梅花鹿行為的分類與定義-------------------------------------------------- ---110 附錄5. 社頂地區穿越線植種相對頻度(%)、鹿、牛及羊攝食情形-------------111 附錄6. 社頂地區三區地表植種及落葉樹種取樣調查之學名及出現數量-----118 附錄7. 社頂地區草食獸食用植物名錄---------------------------------------------- -120 附錄8. 社頂地區台灣梅花鹿所磨植種之名錄-------------------------------------- 128 附錄9. 社頂地區台灣梅花鹿啃剝植種之名錄-------------------------------------- 130

    Akashi, N., and T. Nakashizuka. 1999. Effects of bark stripping by Sika deer (
    Cervus nippon) on population dynamics of a mixed forest in Japan. Forest
    Ecol. & Manage. 113(1): 75-82.
    Anderson, R. C., and A. J. Katz. 1993. Recovery of browse-sensitive tree
    species following release from white-tailed deer Odocoileus virginianus
    Zimmerman browsing pressure. Biological Conservation 63(3): 203-208.
    Armstrong, E., D. Euler, and G. Racey. 1983. Winter bed-site selection by
    white-tailed deer in Central Ontario. J. Wildl. Manage. 47(3): 880-884.
    Asada, M., and K. Ochiai. 1999. Nitrogen content in feces and the diet of Sika
    deer on the Boso Peninsula, central Japan. Ecol. Research 14(3) Sep.: 249-253.
    Augustine, D. J., and L. E. Frelich. 1998. Effects of white-tailed deer on
    populations of an understory forb in fragmented deciduous forests. Conservation
    Biology 12(5) Oct.: 995-1004.
    Beier, P. 1987. Sex differences in quality of white-tailed deer diets. J.
    Mammal. 68:323-329.
    Beier, P., and D. R. Mccullough. 1990. Factors influencing white-tailed deer
    activity patterns and habitatuse. Wildl. Monographs 109:1-51.
    Beringer, J., L. P. Hansen, R. A. Heinen, and N. F. Giessman. 1994. Use of
    dogs to reduce damage by deer to a white pine plantation. Wildl. Soc. Bull. 22
    :627-632.
    Berteaux, D., M. Crete, J. Huot, J. Maltais, and J.P. Ouellet. 1998. Food
    choice by white-tailed deer in relation to protein and energy content of the
    diet: A field experiment. Oecologia 115(1-2) June: 84-92.
    Bertrand, M. R., A. J. Denicola, S. R. Beissinger, and R. K. Swihart. 1996.
    Effects of parturition on home ranges and social affiliations of female white-
    tailed deer. J. Wildl. Manage. 60(4): 899-909.
    Bideau, E., J. F. Gerard, J. P. Vincent, and M. L. Maublanc. 1993. Effects of
    age and sex on space occopation by European roe deer. J. Mammal. 74(3): 745-
    751.
    Bobek, B. 1977. Summer food as the factor limiting roe deer population size.
    Nature 268:47-49.
    Borkowski, J., and K. Furubayashi. 1998. Seasonal changes in number and
    habitat use of foraging Sika deer at the high altitude of Tanzawa Mountains,
    Japan. Acta Theriologica 43(1): 95-106.
    Bowyer, R. T. 1984. Sexual segregation in southern mule deer. J. Mammal. 65:
    410-417.
    Case, R. L., and J. B. Kauffman. 1997. Wild ungulate influences on the
    recovery of  willows, black cottonwood and thin-leaf alder following
    cessation of cattle grazing in northeastern Oregon. Northwest Science 71(2):
    115-126.
    Catt, D. C., and B. W. Staines. 1987. Home range use and habitat selection by
    Red deer (Cervus elaphus) in a Sitka spruce plantation as determined by radio-
    tracking. J. Zool., Lond. 211:681-693.
    Cederlund, G., and H. Sand. 1994. Home range size in relation to age and sex
    in moose. J. Mammal. 75(4): 1005-1012.
    Clutton-Brock, T. H., F. E. Guinness, and S. D. Albon. 1982. Red deer behavior
    and ecology of two sexes. The University of Chicago Press, Chicago, Ill. 378pp.
    Conradt, L., T. H. Clutton-Brock, and F. E. Guinness. 2000. Sex differences in
    weather sensitivity can cause habitat segregation: red deer as an example.
    Animal behaviour 59:1049-1060.
    DeGarmo, W. R., and J. Gill. 1958. West virginia whitetails. Conser. Comm.
    Bull 4 West virginia Div. Game Manage Charleston 87pp.
    Dhungel, S. K., and B. W. O’Gara. 1991. Ecology of the hog deer in Royal
    Chitwan National Park, Nepal. Wildl. Monogr. 119:1-40.
    Eiberle, K., and B. Riedi. 1985. Critical browsing limits for pine Pinus-
    sylvestris in a red deer wintering area. 〔German〕 Zeitschrift Fuer
    Jagdwissenschaft 31(2): 103-116.
    Ekblad, R. L., J. W. Stuth, and M. K. Owens. 1993. Grazing pressure impacts on
    potential foraging competition between Angora goats and white-tailed deer.
    Small Ruminant Research 11(3): 195-208.
    Endo, A., and T. Doi. 1996. Home range of female sika deer Cervus nippon on
    Nozaki Island, the Goto Archipelago. Mammal Study 21:27-35.
    Feldhamer, G. A. 1980. Cervus nippon. Mammal species. The American Soci. of
    Mammalogists 128:1-7.
    Feldhamer, G. A., K. R. Dixon, and J. A. Champan. 1982. Home range and
    movement of Sika deer in Maryland. Zeitschrift saugetierk. 47(5): 311-316.
    Flynn, A. and A. W. Franzmann. 1987. Mineral element studies in North American
    moose. Swed. Wildl. Res. (Suppl.) 1: 289-300.
    Fraser, K. W. 1996. Comparative rumen morphology of sympatric Sika deer (
    Cervus nippon) and red deer (Cervus elaphus scoticus) in the Ahimanawa and
    Kaweka Ranges, central North Island, New Zealand. Oecologia 105(2) Jan.: 160-
    166.
    Fraser, M. D., and I. J. Gordon. 1997. The diet of goats, red deer and South
    American camelids feeding on three contrasting Scottish upland vegetation
    communities. J. Appl. Ecol. 34(3): 668-686.
    Gaillard, J. M., and D. Delorme. 1989. Selection of bedding sites by young roe
    deer (French). Acta Oecologica. Oecologia Generalis 10(4): 411-418.
    Gill, R. M. A. 1992. A review of damage by mammals in North Temperate forests:
    3. Impact on trees and forests. Forestry (Oxford) 65(4): 363-388.
    Gittleman, J. L., and P. H. Harvey. 1982. Carnivore home range size, metabolic
    needs and ecology. Behav. Ecol. and Sociobiology 10:57-63.
    Hadjisterkotis, E. 1996. Food habits of the Cyprus mouflon Ovis gmelini
    ophion. 〔German〕Zeitschrift Fuer Jagdwissenschaft 42(4): 256-263.
    Harestad, A. S., and F. L. Bunnell. 1979. Home range and body weight - A
    reevaluation. Ecol. 60:389-402.
    Hayes, C. L. and P. R. Krausman. 1993. Nocturnal activity of female desert
    mule deer. J. Wildl. Manage. 57(4): 897-904.
    Hayne, D. W. 1949. Calculation of size of home range. J. Mammal. 30:1-18.
    Heitkonig, I. M. A., and N. Owen-Smith. 1998. Seasonal selection of soil types
    and grass swards by roan antelope in a South African savanna. African J. Ecol.
    36(1) March: 57-70.
    Henke, S. E., S. Demarais, and J. A. Pfister. 1988. Digestive capacity and
    diets of white-tailed deer and exotic ruminants. J. Wildl. Manage. 52(4): 595-
    598.
    Heroldov&aacute;, M. 1996. Dietary overlap of three ungulate species in the
    Palava Biosphere Reserve. Forest Ecol. and Manage. 88:139-142.
    Hester, A. J., I. J. Gordon, G. J. Baillie, and E. Tappin. 1999. Foraging
    behaviour of sheep and red deer within natural heather/grass mosaics. J. Appl.
    Ecol. 36(1) Feb.: 133-146.
    Hobbs, N. T. 1987. Fecal indices to dietary quality: a critique. J. Wildl.
    Manage. 51:317-320.
    Hodgman, T. P., and R. T. Bowyer. 1985. Winter use of arboreal lichens
    ascomycetes by white-tailed deer Odocoileus-virginianus in Maine U.S.A.
    Canadian Field-Naturalist 99(3): 313-316.
    Hofmann, R. R. 1988. Morphophysiological evolutionary adaptations of the
    ruminant digestive physiology in ruminants. A. Dobson and M. J. Dobson (eds).
    Cornell University Press. Ithaca, New York P1-19.
    H&ouml;lzenbein, S., and G. Schwede. 1989. Activity and movements of female
    white-tailed deer during the rut. J. Wildl. Manage. 53(1): 219-223.
    Horwood, M.T., and E.H. Masters. 1970. Sika deer. The British Deer Society.
    30pp.
    Huegel, C. N., R. B. Dahlgren, and H. L. Gladfelter. 1986. Bedsite selection
    by white-tailed deer fawns in Iowa. J. Wildl. Manage. 50(3): 474-480.
    Humphreys, G. G., and T. A. Nelson. 1999. Mortality and home range of white-
    tailed deer on Fort Chaffee Military Reservation, Arkansas. Proceedings of the
    Annual Southeastern Deer Study Group 22:27.
    Ito, T. and S. Takatsuki. 1987. The Distribution and seasonal movements of
    Sika deer (Cervus nippon) in the Mt. Goyo Area, Iwate Prefecture. Bull. Of
    Yamagata Univ., Nat. Sci. 11(4) Jan.: 411-430.
    Jayasekara, P., and S. Takatsuki. 2000. Seasonal food habits of a Sika deer
    population in the warm temperate forest of the western most part of Honshu,
    Japan. Ecol. Res. 15(2) Jun.: 153-157.
    Johansson, A., and O. Liberg. 1996. Function aspects of marking behavior by
    male roe deer (Capreolus capreolus). J. Mammal. 77(2): 558-567.
    Johansson, A., O. Liberg, and L. K. Wahlstrom. 1995. Temporal and physical
    characteristics of scraping and rubbing in roe (Capreolus capreolus). J.
    Mammal. 76(1): 123-129.
    Kaji, K., T. Koizumi, and N. Ohtaishi. 1988. Effects of resource limitation on
    the physical and reproductive condition of Sika deer in Nakanoshima Island,
    Hokkaido. Acta Theriol. 33:187-208.
    Kie, J. G., 1996. The effects of cattle grazing on optimal foraging in mule
    deer (Odocoileus hemionus). Forest Ecol. and Manage. 88:131-138.
    Klein, D. R. 1985. Population Ecology: The interaction between deer and their
    food supply P.13-22. In “Biology of deer production”. Bulletin 22. The royal
    society of New Zealand. Wellington.
    Klein, D. R., and F. Schonheyder. 1970. Variation in ruminal levels among some
    cervids. Can. J. Zool. 48:1437-1442.
    Koeller, J. 1990. The occurrence of the Japanese Sika deer Cervus nippon
    nippon Temminck 1838 and the dybowski deer Cervus nippon hortulorum Swinhoe
    1864 in Hungary. Zeitschrift Fuer Jagdwissenschaft. 36(2): 73-82.
    Koga, T., and Y. Ono. 1994. Sexual differenes in foraging behavior of Sika
    deer, Cervus nippon. J. Mammal. 75(1): 129-135.
    Lang, B. K., and J. E. Gates. 1985. Selection of sites for winter night beds
    by white-tailed deer in a hemlock-northern hardwood forest. American Midland
    Naturalist. 113(2): 245-254.
    Langvatn, R., and T. A. Hanley. 1993. Feeding-patch choice by red deer in
    relation to foraging efficiency. An experiment. Oecologia. 95(2): 164-170.
    Latham, J., B. W. Staines, and M. L. Gorman. 1996. The relative densities of
    red (Cervus elaphus) and roe (Capreolus capreolus) deer and their relationship
    in Scottish plantation forests. J. Zool., Lond. 240:285-299.
    Lee, J. H., I. D. Lee, and H. S. Lee. 1990. Studies on the utilization of
    browse by the Sika deer Cervus nippon I. Food habits of the Sika deer. Korean
    J. Anim. Sci. 32(2): 100-108.
    Lee, G. E., G. C. White, R. A. Carrott, R. M. Bartmann, and A. W. Alldredge.
    1985. Accessing accuracy of a radiotelemetry system for estimating animal
    location. J. Wildl. Manage. 49:658-663.
    Leeuwenberg F. J., S. L. Resende, F. H. G. Rodrigues, and M. X. A. Bizerri.
    1997. Home range, activity and habitat use of the Pampas deer Ozotoceros
    bezoarticus L. 1758(Artiodactyla, Cervidae) in the Brazilian Cerrado. Mammalia
    61(4): 487-495.
    Leslie, D. M., and E. E. Starkey. 1985. Fecal indices to dietary quality of
    cervids in old-growth forests. J. Wildl. Manage. 49:142-146.
    Lindstedt, S. L., B. J. Miller, and S. W. Buskirk. 1986. Home range, time, and
    body size in mammals. Ecology 67:413-418.
    Litchfield, T. R. 1988. Relationships among white-tailed deer rubbing,
    scraping, and breeding activities. MAI 26/03, p. 309.
    Livezey, K. B. 1991. Home range, habitat use, disturbance, and mortality of
    Columbian black-tailed deer in Mendocino National Forest. California Fish and
    Game 77(4): 201-209.
    Loft, E. R., J. W. Menke, and T. S. Burton. 1984. Seasonal movements and
    summer habitats of female black-tailed deer. J. Wildl. Manage. 48(4): 1317-
    1325.
    Loft, E. R., J. W. Menke, and J. G. Kie. 1991. Habitat shifts by mule deer:
    The influence of cattle grazing. J. Wildl. Manage. 55(1): 16-26.
    Lovallo, M. J., and E. M. Anderson. 1996. Bobcat movements and home ranges
    relative to roads in Wisconsin. Wildl. Soc. Bull. 24:71-76.
    Lovari, S., and C. S. Jose. 1997. Wood dispersion affects home range size of
    femal roe deer. Behavioural Processes 40:239-241.
    MacArthur, R. H., and E. R. Pianka. 1966. On optimal use of a patchy
    environment. Am. Nat. 100:603-609.
    Mackie, R. J. 1978. Impact of livestock grazing on wild ungulates. Trans.
    North Am. Wildl. Nat. Resour. Conf. 43:462-476.
    Main, M. B., and B. E. Coblenz. 1990. Sexual segregation among ungulates: a
    critique. Wildl. Soc. Bull. 18:204-210.
    Maizeret, C. 1988. Roe deer foraging strategies ecological basis of diet
    diversity. Acta Oecologica Oecologia Applicata. 9(2): 191-212.
    Maizeret, C., and P. Ballon. 1990. Analysis of causal factors behind cervid
    damage on the cluster pine in the lands of Gascony France. Gibier Faune
    Sauvage 7:275-294.
    Mann, J. C. E., and R.J. Putman. 1989. Diet of British sika deer in
    contrasting enviroment. Acta theriol. 34(6): 97-109.
    Manser, M. B., and P. N. M. Brotherton. 1995. Environmental constraints on the
    foraging behaviour of a dwarf antelope (Madoqua kirkii). Oecologia. 102(4):
    404-412.
    Martin, P. 1993. Measuring behavior. University Press, Cambridge. P84-100.
    Maruyama, N. 1981. A study of the seasonal movements and aggregation pattern
    of Sika deer. Bull. Fac. Agric. Tokyo Univ. Agric. Technol. 23, 85pp (in
    Japanese with English summary).
    Maruyama, N., Y. Totake, and R. Okabayashi. 1976. Seasonal movement of sika
    deer in Omote-Nikko, Tochigi Prefecture. J. Mammal. Soc. Japan 5/6: 187-198.
    McCorquodale, S. M. 1993. Winter foraging behavior of elk in the shrub-steppe
    of Washington. J. Wildl. Manage. 57(4): 881-890.
    McCullough, D. R. 1974. Status of larger mammals in Taiwan. Tourism Bureau,
    Taipei, Taiwan, R. O. C. 35pp.
    McCullough, D. R. 1979. The George Reserve deer herd: population ecology of a
    k-selected species. Univ. Michigan Press, Ann Arbor. 271pp.
    Mcinnis, M. L., and M. Vavra. 1986. Summer diets of domestic sheep grazing
    mountain meadows in Northeastern Oregon USA. Northwest Science 60(4): 265-270.
    Mcinnis, M. L., and M. Vavra. 1987. Dietary relationships among feral horses,
    cattle and pronghorn in southeastern Oregon U.S.A. J. Range Manage. 40(1): 60-
    66.
    McNab, B. K. 1963. Bioenergetics and the determination of home range size.
    American Naturalist 47:133-140.
    Miller, K. V., and R. L. Marchinton. 1999. Temporal distribution of rubbing
    and scraping by a high-density white-tailed deer, Odocoileus virginianus
    population in Georgia. Canadian Field-Naturalist. 113(3) July-Sept.: 519-521.
    Miller, K. V., R. L. Marchinton, K. J. Forand, and K. L. Johansen. 1987.
    Dominance, testosterone levels, and scraping activity in a captive herd of
    white-tailed deer. J. Mammal. 68(4): 812-817.
    Milton, K., and M. L. May. 1976. Body weight, diet and home range area in
    primates. Nature 259:459-462.
    Miquelle, D. G., and B. V. Van. 1989. Impact of bark stripping by moose on
    aspen-spruce communities. J. Wildl. Manage. 53(3): 577-586.
    Mitchell, B., and D. McCowan. 1986. Patterns of damage in relation to the site
    preferences of deer in an enclosed plantation of sitka spruce and lodgepole
    pine. Scottish Forestry 40(2): 107-117.
    Mitchell, J. E., and R. T. Rodgers. 1985. Food habits and distribution of
    cattle on a forest and pasture range in Northern Idaho USA. J. Range Manage.
    38(3): 214-220.
    Miura, S. 1976. Ecological studies on sika deer in Nara Park with reference to
    spatial structure. Ann. Rep. Nara Deer Res. Assoc. 2:47-61(in Japanese with
    English summary).
    Miura, S. 1984. Social behavior and territoriality in male sika deer (Cervus
    nippon Temmminck, 1838) during the rut. Z. Tierpsychol. 64:33-73.
    Moen, A. N. 1978. Seasonal changes in heart rates, activity, metabolism, and
    forage intake of white-tailed deer. J. Wildl. Manage. 42:715-738.
    Montgomery, G. G. 1963. Nocturnal movements and activity rhythms of white-
    tailed deer. J. wildl. Manage. 27(3): 422-427.
    Morimura, N., and Y. Ueno. 1998. Behavior patterns of 9 nammals in the zoo:
    The comparison among species, and different environments. Japanese J. Anim.
    Psychology. 48(1) June: 33-45.
    Motta, R. 1996. Impact of wild ungulates on forest regeneration and tree
    composition of mountain forests in the Western Italian Alps. Forest Ecol. and
    Manage. 88(1-2): 93-98.
    Motta, R., and P. Nola. 1996. Fraying damages in the subalpine forest of
    paneveggio (Trento, Italy): A dendroecological approach. Forest Ecol. and
    Manage. 88(1-2): 81-86.
    Mould, E. D., and C. T. Robbins. 1981. Nitrogen metabolism in elk. J. Wildl.
    Manage. 45:323-334.
    Mrlik, V. 1988. Daily activity and space use of male Sika deer Cervus-nippon
    Dybowskii during the rut. Folia Zool. 37(4): 319-332.
    Murden, S. B., and K. L. Risenhoover. 1993. Effects of habitat enrichment on
    patterns of diet selection. Ecol. Applications 3(3): 497-505.
    Mysterud, A. 1999. Seasonal migration pattern and home range of roe deer (
    Capreolus capreolus) in an altitudinal gradient in southern Norway. J. Zool.,
    Lond. 247:479-486.
    Mysterud, A., and P. K. Larsen, R. A. Ims, and E. Ostbye. 1999. Habitat
    selection by roe deer and sheep: Does habitat ranking reflect resource
    availability﹖Can. J. Zool. 77(5) May : 776-783.
    Mysterud, A., and E. Ostbye. 1995. Bed-site selection by European roe deer (
    Capreolus capreolus) in southern Norway during winter. Can. J. Zool. 73(5):
    924-932.
    Mysterud, A., and I. Mysterud. 2000. Ecological effects of domestic ruminant 
    grazing: Interactions between large herbivores. 〔Norwegian〕 Fauna (Ooslo).
    〔print〕53(1): 22-51.
    Nams, V. O. 1990. Locate II vser's guide. Pacer Computer Software. Truro, N.
    S.
    Obrtel, R., V. Holisova, and I. Kozena. 1985. The winter diet of Sika deer (
    Cervus nippon) in the Bouzovsko area Czechoslovakia. Folia Zool. 34(1): 1-22.
    Ockenfels, R. A. and D. E. Brooks. 1994. Summer diurnal bed sites of coues
    white-tailed deer. J. Wildl. Manage. 58(1): 70-75.
    Odajima, M., Y. Kajita, K. T. Nam, S. R. Lee, H. Senge, K. Katoh, Y. Shoji, M.
    Ohta, and Y. Sasaki. 1991. Seasonal changes in food passage and digestibility
    in Japanese Sika deer Cervus nippon and sheep under restricted feeding
    condition. Animal Science & Technology 62(3): 308-313.
    Opperman, J. J., and A. M. Merenlender. 2000. Deer Herbivory as an ecological
    constraint to restoration of degraded riparian corridors. Restoration Ecol. 8(
    1) March : 41-47.
    Padmalal, U. K. G. K., and S. Takatsuki. 1994. Age-sex differences in the
    diets of Sika deer on Kinkazan Island, northern Japan. Ecol. Research 9:251-
    256.
    Palat, M., F. Vasicek, V. Henzlik, and H.D. Kasperidus. 1994. Condition of
    damage to Norway spruce stands in the Czech Republic. Lesnictvi (Prague). 40(
    6):217-237.
    Parks C. G., L. Bednar, and A. R. Tiedemann. 1998. Browsing ungulates: An
    important consideration in dieback and mortality of Pacific yew (Taxus
    brevifolia) in a northeastern Oregon stand. Northwest Science 72(3) Aug.: 190-
    197.
    Putman, R. J. 1986. Competition and coexistence in a multi-species grazing
    system. Acta Theriologica 13(21): 271-292.
    Ram&iacute;rez, R. G., J. B. Quintanilla, and J. Aranda. 1997. White-tailed
    deer food habits in northeastern Mexico. Small Ruminant Research 25:141-146.
    Renecker, L. A., and R. J. Hudson. 1989. Seasonal activity budgets of moose in
    Aspen-dominated boreal forests. J. Wildl. Manage. 53(2): 296-302.
    Robbins, C. T. 1992. Wildlife feeding and nutrition. Academic Press, Inc. San
    Diego. C.A. 352pp.
    Schmitz, O. J. 1991. Thermal constraints and optimization of winter feeding
    and habitat choice in white-tailed deer. Holarctic Ecol. 14(2): 104-111.
    Schoen, J. W., and M. D. Kirchhoff. 1985. Seasonal distribution and home range
    patterns of Sitka black-tailed deer on Admiralty Island, southeast Alaska. J.
    Wildl. Manage. 49(1): 96-103.
    Schoener, T. W. 1968. Sizes of feeding territories among birds. Ecology 49:
    123-141.
    Schwartz, C. C. 1992. Physiological and nutritional adaptations of moose to
    northern environments. Alces (Suppl.) 1:139-155.
    Scott, D. 1998. Impact of red deer on a Scots pine plantation after removal of
    deer fencing. Scottish Forestry 52(1): 8-13.
    Segelquist, C. A., H. L. Short, F. D. Ward, and R. G. Leonard. 1972. Quality
    of some winter deer forages in the Arkansas Ozarks. J. Wildl. Manage. 36:174-
    177.
    Seip, D. R., and F. L. Bunnell. 1985. Foraging behavior and food habits of
    stones sheep Ovis dalli stonei. Can. J. Zool. 63(7): 1638-1646.
    Sekine, T., and H. Sato. 1992. Tree barking by Sika deer Cervus nippon
    Temminck, on Mt. Odaigahara. Japanese J. Ecol. (Sendai) 42(3): 241-248.
    Severson, K. E., and A. L. Medina. 1983. Deer and elk habitat management in
    the Southwest. J. Range Manage. Monogr. 2:1-64.
    Shigematsu, Y., K. Ochiai, and M. Asada. 1994. Animal tracking by radio-
    telemetry. In Reports of management for Sika deer of Boso peninsula in Chiba
    prefecture, II. Chiba prefecture, 59pp (in Japanese).
    Sinclair, A. R. E., C. J. Krebs, and J. N. M. Smith. 1982. Diet quality and
    local limitation in herbivores: the case of the snowshoe hare. Can. J. Zool.,
    60:889-897.
    Song, Y. 1993. Diurnal activity rhythms of Eld's deer on Hainan Island, China.
    Eiserier Science Publishers B. V., Deer of China, 214-219.
    Staines, B. W., and J. M. Crisp. 1978. Observations on food quality in
    Scottish Red deer (Cervus elaphus) as determined by chemical analysis of the
    rumen contents. J. Zool. Lond. 185:253-277.
    Storm, G. L., D. F. Cottam, and R. H. Yahner. 1995. Movements and habitat use
    by female deer in historic areas at Gettysburg, Pennsylvania. Northeast
    Wildlife 52:49-57.
    Suda, K. 1997. Rumen contents and food selectivity of Sika deer (Cervus
    nippon) on Tsushima Islands. Wildl. Conservation Japan 2(3): 125-134.
    Sweeney, J. R. 1970. The effects of harassment by hunting dogs on the movement
    patterns of white-tailed deer on the Savannah River Plant, South Carolina.
    Thesis, University of Georgia, Athens, U.S.A.
    Swihart, R. K., N. A. Slade, and B. J. Bergstrom. 1988. Relating body size to
    the rate of home range use in mammals. Ecol. 69:393-399.
    Swihart, R. K., P. M. Picone, A. J. DeNicola, and L. Cornicelli. 1995. Ecology
    of urban and suburban white-tailed deer. Pages 35-44 in J. B. McAninch, editor.
    Takahasi, H., and K. Kaji. 2001. Fallen leaves and unpalatable plants as
    alternative foods for sika deer under food limitation. Ecological Resaerch. 16(
    2): 257-262.
    Takatsuki, S. 1980. Food habits of Sika deer on Kinkazan Island. Sci. Rep.
    Tohoku Uiv. Ser. IV (Biol.). 38(1): 7-31.
    Takatsuki, S. 1982. Ecological studies on effect of Sika deer (Cervus nippon)
    on vegetation Ⅲ. The vegetation of Iyo-Kashima Island, southwestern Shikoku,
    with reference to grazing effect of Sika deer. Ecol. review 20(1): 15-29.
    Takatsuki, S. 1983. The importance of Sasa nipponica as a forage for Sika
    deer (Cervus nippon) in Omote-Nikko. Japan J. of Ecol. 33:17-25.
    Takatsuki, S. 1986. Food habits of Sika deer on Mount Goyo northern Honshu
    Japan. Ecol. Research 1(2): 119-128.
    Takatsuki, S. 1988. The weight contributions of stomach compartments of Sika
    deer. J. Wildl. Manage. 52(2): 313-316.
    Takatsuki, S. 1990. Summer dietary compositions of Sika deer on Yakushima
    Island, Southern Japan. Ecol. Research. 5:253-260.
    Takatsuki, S., H. Kawahara, and C. Torisu. 1984. Fecal analysis of Sika deer
    on Nozaki Island, the Goto Island, northwestern Kyushu. The Bulletin of
    Nagasaki Institute of Applied Science 25(1): 37-44.
    Tatsuhara, A., and Y. Fujimaki. 1987. Food habit analysis by using opal
    phytoliths in Cervus nippon yesoensis. Research Bulletin of Obihiro University
    Series I 15(2): 85-90.
    Telfer, E. S. 1994. Cattle and cervid interactions on a foothills watershed in
    southwestern Alberta. Canadian Field-Naturalist 108(2): 186-194.
    Thill, R. E., and A. Jr. Martin. 1989. Deer and Cattle diets on heavily grazed
    pine-bluestem range. J. Wildl. Manage. 53(3): 540-548.
    Thomas, J. W., and D. E. Toweill. 1982. Elk of North America-ecology and
    management. Wildlife Management Institude Stackpole books. 698pp.
    Tierson, W. C., G. F. Mattfeld, R. W. Sage, and D. F. Behrend. 1985. Seasonal
    movements and home ranges of white-tailed deer in the Adirondack. J. Wildl.
    Manage. 49(3): 760-769.
    Tixier, H., C. Maizeret, P. Duncan, R. Bertrand, C. Poirel, and M. Roger.
    1998. Development of feeding selectivity in roe deer. Behavioural Processes 43:
    33-42.
    Ullrey, D. E. 1982. Nutrition and antler development in white-tailed deer.
    Pages 49-59 in Brown, R. D. ed. Antler development in Cervidae, Caesar Kleberg
    Wildlife Research Institute, Texas A&Z University, Kingsville, Texas.
    Uresk, D. W. 1986. Food habits of cattle on mixed-grass prairie on the
    northern Great Plains USA. Prairie Naturalist 18(4): 211-218.
    Van, W. S. E. 1995. The potential role of large herbivores in nature
    conservation and extensive land use in Europe. Biological Journal of the
    Linnean Society 56(SUPPL. A): 11-23.
    Veen, H. E. Van de. 1979. Food selection and habitat use in the red deer (
    Cervus elaphus L.) PhD thesis. University of Groningen.
    Vincent, J. P., E. Bideau, A. J. M. Hewison, and J. M. Angibault. 1995. The
    influence of increasing density on body weight, kid production, home range and
    winter grouping in roe deer (Capreolus capreolus). J. Zool., Lond. 236:371-382.
    Voelk, Von F. H. 1999. Bark peeling frequency in the alpine provinces of
    Austria: The importance of forest structure and red deer management.
    Zeitschrift Fuer Jagdwissenschaft 45(1): 1-16.
    Vogel, W. O. 1989. Response of deer to density and distribution of housing in
    Montana. Wildl. Soc. Bull. 17:406-413.
    Wahlstr&ouml;m, L. K., and O. Liberg. 1995. Patterns of dispersal and seasonal
    migration in roe deer (Capreolus capreolus). J. Zool., Lond. 235:455-467.
    Wallmo, O. C. and D. J. Neff. 1970. Direct observation of tamed deer to
    measure their consumption of natural forage. In Range and wildlife habitat
    evaluation a research symposium, pp. 105-110. Misc. publ. 1147. Washington, D.
    C.: USDA For. Serv.
    Watanabe, T., and S. Takatsuki. 1993. Comparison of nitrogen and fiber
    concentrations in rumen and fecal contents of Sika deer. J. Mamm. Soc. Japan
    18(1): 43-48.
    Weckerly, F. W. 1993. Intersexual resource partitioning in Black-tailed deer:
    a test of body size hypothesis. J. Wildl. Manage. 57(3): 475-494.
    Weerasinghe, U. R., and S. Takatsuki. 1999. A record of acorn eating by Sika
    deer in western Japan. Ecological Research 14(2) Jun.: 205-209.
    Welch, D., and D. Scott. 1998. Bark-striping damage by red deer in a Sitka
    spruce forest in western Scotland: IV. Survival and performance of wounded
    trees. Forestry (Oxford) 71(3): 224-235.
    Whitehead, G. K. 1972. Deer of the worid. Constable and Company Ltd., London.
    194pp.
    Wikeem, B. M., and M. D. Pitt. 1992. Diet of California bighorn sheep, Ovis
    canadensis californiana, in British Columbia: Assessing optimal foraging
    habitat. Canadian Field-Naturalist 106(3): 327-335.
    Wofford, H., J. L. Holechek, M. L. Galyean, J. D. Wallace, and M. Cardenas.
    1985. Evaluation of fecal indices to predict cattle quality. J. Range Manage.
    38:450-454.
    Wood, A. K. 1988. Use of shelter by mule deer during winter. Prairie
    Naturalist 20:15-22.
    Wright, I. A., and J. A. Milne. 1996. Aversion of red deer and roe deer to
    denatonium benzoate in the diet. Forestry (Oxford) 69(1): 1-4.
    Yabe, T. 1994. Management of Wildlife Habitat. Ph. D. thesis Fac. Agr.
    Hokkaido Univ., 123pp (in Japanese).
    Yokoyama, S., T. Koizumi, and E. Shibata. 1996. Food habits of Sika deer as
    Assessed by fecal analysis in Mt. Ohdaigahara, central Japan. J. of Forest
    Research 1:161-164.
    Yokoyama, S., I. Maeji, T. Ueda, M. Ando, and E. Shibata. 2001. Impact of
    bark stripping by Sika deer, Cervus nippon, on subalpine coniferous forests in
    central Japan. Forest Ecol. and Manage. 140(2-3) 15 January: 93-99.
    Zejda, J., M. Rebickova, and M. Homolka. 1985. Study of behavior in field roe
    deer Capreolus capreolus. Prirodovedne Prace Ustavu Ceskoslovenske Akademie
    Ved V Brne. 19(12): 1-37.
    Zhang, E. 2000. Daytime activity budgets of the Chinese water deer. Mammalia
    64(2): 163-172.
    中央氣象局月報,1995年6月至1999年6月。
    中村孝志。1956。十七世紀臺灣鹿皮之出產及其對日貿易。臺灣銀行季刊10(2): 131-147
    頁。
    中國科學院植物研究所。1987。中國高等植物科屬檢索表。科學出版社。733頁。
    王穎。1985。台灣梅花鹿之行為研究。台灣梅花鹿復育之研究七十三年度報告。內政部營
    建署墾丁國家公園管理處。保育研究報告第18號。102-179頁。
    王穎和印莉敏。1992。墾丁國家公園有蹄類及食肉目動物調查。內政部營建署墾丁國家公
    園管理處。保育研究報告第80號。54頁。
    王穎, 郭國偉和楊美玉。1993。台灣地區梅花鹿原野棲地調查及野放可行性之評估內政部
    營建署墾丁國家公園管理處。保育研究報告第88號。60頁。
    王穎, 陳順其, 林政田, 詹世琛和張容瑱。1997. 墾丁國家公園台灣梅花鹿野放後之族群
    監測及生態環境影響研究。內政部營建署墾丁國家公園管理處。保育研究報告第91號。45
    頁。
    王穎, 陳順其, 孫元勳, 林政田和廖秀芬。1996. 墾丁國家公園台灣梅花鹿野放後之生態
    學研究。內政部營建署墾丁國家公園管理處。保育研究報告第93號。63頁。
    王穎, 陳輝勝, 吳幸如和蘇銘言。1992。台灣梅花鹿行為及棲地利用之研究。內政部營建
    署墾丁國家公園管理處。保育研究報告第78號。41頁。
    王穎, 詹世琛, 陳順其和陳芬蕙。1998. 墾丁國家公園台灣梅花鹿野放後族群之監測及生
    態環境影響評估。內政部營建署墾丁國家公園管理處。保育研究報告第100號。65頁。
    江樹生。1985。梅花鹿與台灣早期歷史關係之研究。台灣梅花鹿復育之研究七十三年度報
    告。內政部營建署墾丁國家公園管理處。3-62頁。
    宋延齡。1997。兩種食性研究方法在海南坡鹿食性研究中的比較。獸類學報 17(2): 151-
    154頁。
    施宗雄, 楊錫坤, 宋尚美和黃國雄。1985。台灣梅花鹿養殖現況調查研究。台灣梅花鹿復
    育之研究七十三年度報告。墾丁國家公園管理處。248-275頁。
    胡正恆和王穎。1994. 社頂地區台灣梅花鹿之食性初探。師大生物學報 29(1): 11-20頁。
    兼本 正, 米須 清智。1996。行動圈及與日周活動。梅花鹿 Cervus nippon keramae 保
    護對策緊急實態調查報告書。沖繩縣教育委員會。175-188頁。
    高槻成紀, 鈴木(K. Suzuki)。1982。日本中部丹澤山梅花鹿冬季期間胃內食物分析。日
    本 Tohoku 大學。
    夏良宙, 王穎, 程中江和陳寶忠。1990。台灣梅花鹿攝食喜好性試驗。台灣梅花鹿復育之
    研究七十八年度報告。內政部營建署墾丁國家公園管理處。49-73頁。
    秦榮前, 范玉琳, 梁鳳錫, 何春林和李尚仁。1994。中國梅花鹿。中國農業出版社。292頁

    盛和林, 大泰司紀之, 張恩迪, 查普曼, 曹克清, 李文軍, 馬逸清和徐宏發。1992。中國
    鹿類動物。華東師範大學出版社。305頁。
    郭延蜀, 胡錦矗, 羅代華, 色科, 任樹培和鄒洪富。1991。四川梅花鹿的社群行為研究。
    獸類學報 11(3):165-170頁。
    郭國偉。1994。台灣梅花鹿野外行為及棲地利用之研究。國立台灣師範大學生物研究所碩
    士論文。42頁。
    陳雲倩。1989。社頂植群分析及牛羊放牧影響之研究。國立台灣大學森林研究所碩士論文
    。101頁。
    陳翼勝, 鄭碩和吳征鎰。1997。中國有毒植物。地景企業股份有限公司。827頁。
    張慶恩, 葉慶龍和鍾玉龍。1985。墾丁國家公園社頂自然公園植被及景觀調查規劃報告。
    內政部營建署墾丁國家公園管理處。
    楊錫坤和馬春祥。1994。臺灣梅花鹿之生殖性狀與鹿角週期。中國畜牧學會會誌 23(1):
    43-55頁。
    裴家騏和李佩珍。1999。梅花鹿對綠島外緣開闊草生地的利用。中華林學季刊 32(4):
    425-440頁。
    劉和義。1992。綠島野放梅花鹿生態之調查研究。觀光局東部海岸風景區管理處。
    劉棠瑞和劉儒淵。1977。恆春半島南仁山區植群生態與植物區系之研究。省立博物館科學
    年刊 20:51-150頁。
    潘明雄。個人通訊。潘明雄先生為臺灣梅花鹿復育區管理人員。
    蘇鴻傑。1985。台灣梅花鹿天然生育地之植群分析及其在墾丁國家公園內復育地之勘選。
    台灣梅花鹿復育之研究七十三年度報告。墾丁國家公園管理處。63-99頁。
    蘇鴻傑和陳雲倩。1990。台灣梅花鹿對社頂地區植群影響之研究。台灣梅花鹿復育之研究
    七十八年度報告。墾丁國家公園管理處。18-48頁。
    蘇鴻傑和楊勝任。1988。墾丁國家公園社頂地區之植群生態與演替。台灣梅花鹿復育研討
    會專輯。內政部營建署墾丁國家公園管理處。141-163頁。
    蘇鴻傑, 楊勝任和陳雲倩。1989。台灣梅花鹿對社頂地區植群影響效益之研究—社頂地區
    之植群生態與演替。台灣梅花鹿復育之研究七十七年度報告。墾丁國家公園管理處。42-69
    頁。

    QR CODE